Fishery Rresources and Environment Key Field Station of Upper-middle Reaches of the Yangtze River, Yangtze River Fisheries Research Institute, Chinese Academy of Fishery Sciences, Ministry of Agriculture, Wuhan 430223,China

Fisheries Research Institute of Jiangxi Province, Nanchang 330039, China

农业部长江中上游渔业资源环境重点野外科学观测试验站,中国水产科学研究院长江水产研究所,武汉430223

江西省水产科学研究所,南昌330039