Administrative Authorities of Huanglian Mountains National Nature Reserve, Luchun, Yunnan 662500

云南黄连山国家级自然保护区管理局,云南绿春662500

State Key Laboratory of Systematic and Evolutionary Botany, Chinese Academy of Sciences, Beijing 100093

中国科学院植物研究所系统与进化植物学国家重点实验室,北京 100093