Key Laboratory of Petroleum Geochemistry,Research Institute of Petroleum Exploration and Development,Beijing 100083,China

Key Laboratory of Deep Petroleum Intelligent Exploration and Development,Institute of Geology and Geophysics,Chinese Academy of Sciences,Beijing 100029,China

National Key Laboratory for Multi-Resource Collaborated Green Development of Continental Shale Oil,Daqing 163712,China

National Key Laboratory of Petroleum Resources and Engineering,Research Centre for Geomicrobial Resources and Application,Unconventional Petroleum Research Institute,College of Geosciences,China University of Petroleum,Beijing 102249,China