Ecologie Systématique Evolution,Université Paris-Sud,CNRS,AgroParisTech,Université Paris-Saclay,Orsay Cedex F-91405,France

Ministry of Education,Key Laboratory for Biodiversity Science and Ecological Engineering,College of Life Sciences,Beijing Normal University,Beijing 100875,China

Chasséstraat 34-2,Amsterdam NL-1057JG,The Netherlands

Department—Biology and Environment and Climate Change Canada,University of Western Ontario,Room 2025 BGS Building,1151 Richmond Street,London,ON,Canada N6A 5B7