State Forestry Administration of China Key Laboratory for Biodiversity Conservation in Mountainous Areas of Southwest Karst, School of Life Sciences, Guizhou Normal University, Guiyang, Guizhou 550001, China
Ministry of Education Key Laboratory for Ecology of Tropical Islands, College of Life Sciences, Hainan Normal University, Haikou, Hainan 571158, China
This work was supported by the National Natural Science Foundation of China).L.W.W.was funded by the Initial Fund Key Laboratories of Guizhou Province (2011-4005) and Joint Fund of the Natio