College of Life Sciences and Technology,Central South University of Forestry and Technology,Changsha,Hunan 410004,China

CAS Key Laboratory of Animal Ecology and Conservation Biology,Institute of Zoology,Beijing 100101,China

Shaanxi Key Laboratory for Animal Conservation,College of Life Sciences,Northwest University,Xi'an,Shaanxi 710069,China

Key Laboratory of Southwest China Wildlife Resources Conservation(Ministry of Education)China West Normal University,Nanchong,Sichuan 637009,China

College of Ecology,Lanzhou University,Lanzhou,Gansu 730000,China

State Key Laboratory of Environmental Criteria and Risk Assessment,Chinese Research Academy of Environmental Sciences,Beijing 100012,China

College of Life Sciences,University of Chinese Academy of Sciences,Beijing 100101,China

Center for Excellence in Animal Evolution and Genetics,Chinese Academy of Sciences,Kunming,Yunnan 650223,China