Key Laboratory of Freshwater Fish Reproduction and Development(Ministry of Education),Southwest University School of Life Sciences,Chongqing 400715,China
College of Fisheries,Southwest University,Rongchang,Chongqing 402460,China
Academy of Plateau Science and Sustainability,Qinghai Normal University,Xining,Qinghai 810008,China
National Natural Science Foundation of ChinaFundamental Research Funds for the Central Universities