CAS Key Laboratory of Animal Ecology and Conservation Biology,Institute of Zoology,Chinese Academy of Sciences,Beijing 100101,China

College of Life Sciences,University of Chinese Academy of Sciences,Beijing,100049,China

School of Ecology and Nature Conservation,Beijing Forestry University,Beijing,100083,China

College of Life Sciences and Technology,Central South University of Forestry and Technology,Changsha,Hunan 410004,China

College of Forestry,Central South University of Forestry and Technology,Changsha,Hunan 410004,China

Center for Excellence in Animal Evolution and Genetics,Chinese Academy of Sciences,Kunming,Yunnan 650223,China