1. Department of Vertebrate Zoology,Lomonosov Moscow State University,Moscow 119991,Russia
2. Joint Vietnam-Russia Tropical Science and Technology Research Centre,Cau Giay,Hanoi 122000,Vietnam;A.N.Severtsov Institute of Ecology and Evolution,Russian Academy of Sciences,Moscow 119071,Russia
3. Joint Vietnam-Russia Tropical Science and Technology Research Centre,Cau Giay,Hanoi 122000,Vietnam
4. Department of Vertebrate Zoology,Lomonosov Moscow State University,Moscow 119991,Russia;Joint Vietnam-Russia Tropical Science and Technology Research Centre,Cau Giay,Hanoi 122000,Vietnam