Guangdong Provincial Key Laboratory of Plant Adaptation and Molecular Design,Innovative Center of Molecular Genetics and Evolution,School of Life Sciences,Guangzhou University,Guangzhou 510006,China

College of Agronomy,China Agricultural University,Beijing 100193,China

College of Life Sciences,Fujian Agriculture and Forestry University,Fuzhou,Fujian 350002,China

Synthetic and Functional Biomolecules Center,Beijing National Laboratory for Molecular Sciences,Key Laboratory of Bioorganic Chemistry and Molecular Engineering of Ministry of Education,College of Chemistry and Molecular Engineering,Peking University,Beijing 100871,China

Kingagroot Co.,Ltd.,Qingdao,Shandong 266000,China

Tsinghua University-Peking University Joint Center for Life Sciences,Beijing 100871,China