JSPS KAKENHIJSPS KAKENHIMinistry of Education,Culture,Sports,Science and Technology of JapanCrossministerial Moonshot Agriculture,Forestry and Fisheries Research and Development Program grant from the Cabinet Office,Biooriented Technology Research Advancement InstitutionPriority Research Project(foR-F)of Fukushima Universitycommissioned research fund provided by F-REI