Department of Human Oncology,University of Wisconsin School of Medicine and Public Health,University of Wisconsin,Madison,WI 53705,USA

Department of Biomedical Engineering,University of Wisconsin-Madison,Madison,WI 53705,USA

University of Wisconsin Carbone Cancer Center,Madison,WI 53705,USA

Department of Biostatistics and Medical Informatics,University of Wisconsin-Madison,Madison,WI 53706,USA

Department of Biology,Tufts University,Medford,MA 02155,USA

Department of Pathology and Laboratory Medicine,University of Wisconsin School of Medicine and Public Health,University of Wisconsin,Madison,WI 53705,USA