SXMU-Tsinghua Collaborative Innovation Center for Frontier Medicine,Taiyuan 030001,China

Institute for Immunology,Department of Basic Medical Sciences,School of Medicine,Tsinghua University,Beijing 100084,China

Tsinghua-Peking Center for Life Sciences,Tsinghua University,Beijing 100084,China

Shanxi Medical University,Taiyuan 030012,China

Department of Surgery,First Hospital of Shanxi Medical University,Shanxi Provincial Key Laboratory for Digestive Diseases and Organ Transplantation,Taiyuan 030001,China

Beijing Key Laboratory for Immunological Research on Chronic Diseases,Tsinghua University,Beijing 100084,China