国家重点研发计划国家重点研发计划国家重点研发计划国家重点研发计划国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金国家自然科学基金NSFC/RGC Joint Grant Scheme(2022、2023)中国科学院战略规划重点项目Key Research Program,CASInternational Cooperation Program,CASCAS Project for Young Scientists in Basic ResearchGuangdong Province Science and Technology ProgramGuangdong Province Science and Technology ProgramGuangdong Province Science and Technology ProgramGuangdong Province Science and Technology ProgramGuangdong Province Science and Technology ProgramGuangdong Province Science and Technology ProgramGuangdong Province Science and Technology Program广州市科技计划广州市科技计划广州市科技计划广州市科技计划广州市科技计划HealthInnoHK funding support from the Innovation Technology Commission of the Hong Kong SAR,Basic Research Project of Guangzhou Institutes of Biomedicine and HealthChinese Academy of Sciences and CAS Youth Innovation Promotion Association