State Key Laboratory of Soil and Sustainable Agriculture,Changshu National Agro-Ecosystem Observation and Research Station,Institute of Soil Science,Chinese Academy of Sciences,Nanjing 210008,China

University of Chinese Academy of Sciences,Nanjing 211135,China

University of Chinese Academy of Sciences,Beijing 100049,China

Department of Plant Sciences,University of California,Davis CA 95616,USA

Department of Agronomy,Iowa State University,Ames IA 50011,USA

National Engineering Research Center for Wheat,State Key Laboratory of Wheat and Maize Crop Science,Henan Agricultural University,Zhengzhou 450046,China

Jiangsu Key Laboratory of Low Carbon Agriculture and GHGs Mitigation,College of Resources and Environmental Sciences,Nanjing Agricultural University,Nanjing 210095,China

College of Environmental and Resource Sciences,Zhejiang University,Hangzhou 310058,China