Key Laboratory of Animal Ecology and Conservation Biology,Institute of Zoology,Chinese Academy of Sciences,Beijing 100101,China

University of Chinese Academy of Sciences,Beijing 100049,China

Arctic Seasonal Timekeeping Initiative(ASTI),Department of Arctic and Marine Biology UiT,The Arctic University of Norway,Tromsø 9037,Norway

Cardiff University-Institute of Zoology Joint Laboratory for Biocomplexity Research,Chinese Academy of Sciences,Beijing 100101,China

Center for Excellence in Animal Evolution and Genetics,Chinese Academy of Sciences,Kunming 650223,China