State Key Laboratory of Tibetan Plateau Earth System,Resources and Environment(TPESER),Institute of Tibetan Plateau Research,Chinese Academy of Sciences,Beijing 100101,China

Palaeo-Research Institute,University of Johannesburg,Auckland Park 2006,South Africa

Key Laboratory of Vertebrate Evolution and Human Origins,Institute of Vertebrate Paleontology and Paleoanthropology,Chinese Academy of Sciences,Beijing 100044,China

Department of Archaeology,University of Exeter,Exeter EX4 4QE,UK

School of Geography,Archaeology and Environmental Studies,University of the Witwatersrand,Johannesburg WITS 2050,South Africa