Center for Cell Structure and Function,Shandong Provincial Key Laboratory of Animal Resistance Biology,Haihe Laboratory of Cell Ecosystem,Collaborative Innovation Center of Cell Biology in Universities of Shandong,College of Life Sciences,Shandong Normal University,Jinan 250014,China

State Key Laboratory of Medicinal Chemical Biology,College of Life Sciences,Nankai University,Tianjin 300071,China

Department of Hematology,Tianjin Medical University General Hospital,Tianjin 300052,China

Department of Cell Biology,School of Basic Medical Sciences,Cheeloo College of Medicine,Shandong University,Jinan 250012,China

Laboratory of Tissue Homeostasis,Haihe Laboratory of Cell Ecosystem,Tianjin 300462,China