Group of Alpine Paleoecology and Human Adaptation(ALPHA),Institute of Tibetan Plateau Research,Chinese Academy of Sciences,Beijing 100101,China

State Key Laboratory of Tibetan Plateau Earth System,Environment and Resources(TPESER),Institute of Tibetan Plateau Research,Chinese Academy of Sciences,Beijing 100101,China

University of Chinese Academy of Sciences,Beijing 100049,China

School of History and Culture,Lanzhou University,Lanzhou 730000,China

Key Laboratory of Western China s Environmental Systems(Ministry of Education),College of Earth and Environmental Sciences,Lanzhou University,Lanzhou 730000,China

School of Environmental Studies, China University of Geosciences, Wuhan 430078, China

State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Wuhan 430078, China

Yazhouwan National Laboratory,Sanya 572024,China

Department of Anthropology,Washington University in St.Louis,St.Louis MO 63130,USA

Key Laboratory of Animal Genetics,Breeding and Reproduction of Shaanxi Province,College of Animal Science and Technology,Northwest A&F University,Yangling 712100,China