The Behavior Ethogram and PAE Coding System of Wild Yangtze Finless Porpoise in Nanjing
The behavior ethogram is a behavior catalog compiled based on the identification and classification of animal behaviors,which is an important foundation and scientific basis for further research on animal behavior ecology.From January 2019 to December 2023,112 d vessel-based survey and 10 d land-based drone monitoring in Nanjing Yangtze Finless Porpoise provincial nature reserve were conducted.The study recorded 4 258 times of porpoises'surfacing,and constructed behavior ethogram and PAE(Posture-Act-Environment)coding system.13 P codes,26 A codes and 10 E codes were defined and formed 61 behaviors,all these were divided into 10 general categories,swimming mode,physiological function behavior,fin limb behavior,body behavior,jumping behavior,social behavior,mother child dolphin behavior,predation behavior,mating behavior and spiting behavior.Comparing with porpoise in the semi natural waters of Tongling,47 behaviors were same,most of which were necessary for the basic life activities.14 behaviors occurred only in nature habitat of Nanjing,that may be adaptive strategies for busy shipping traffic,41 behaviors,such as conflict,sliding,and hanging,were only found in Tongling,that might be related to artificial feeding in semi natural waters.There were significant differences in the frequency of some behaviors occurring during dry and flood seasons,as well as under different wind conditions,which might be related to parenting and environmental adaptability.The present study constructed the behavior ethogram and first PAE coding system of wild Yangtze Finless Porpoise in China,providing important reference for species protection and management.