State Key Laboratory of Genetic Engineering at School of Life Sciences,Human Phenome Institute,Obstetrics and Gynecology Hospital,Fudan University,Shanghai 200438,China

Obstetrics and Gynecology Hospital,Institute of Reproduction and Development,State Key Laboratory of Genetic Engineering at School of Life Sciences,Fudan University,Shanghai 200011,China

Human Phenome Institute,Zhangjiang Fudan Intemational Innovation Center,Fudan University,Shanghai 201203,China

Obstetrics and Gynecology Hospital,Institute of Reproduction and Development,Fudan University,Shanghai 200011,China

State Key Laboratory of Genetic Engineering at School of Life Sciences,Obstetrics and Gynecology Hospital,Fudan University,Shanghai 200438,China

National Demonstration Center for Experimental Biology Education,School of Life Sciences,Fudan University,Shanghai 200433,China