Center for Cell Structure and Function,Shandong Provincial Key Laboratory of Animal Resistance Biology,Collaborative Innovation Center of Cell Biology in Universities of Shandong,College of Life Sciences,Shandong Normal University,Jinan,Shandong 250014,China

State Key Laboratory of Medicinal Chemical Biology,Haihe Laboratory of Cell Ecosystem,College of Life Sciences,Nankai University,Tianjin 300071,China