Institute of Neuroscience,State Key Laboratory of Neuroscience,CAS Center for Excellence in Brain Science and Intelligence Technology,Chinese Academy of Sciences,Shanghai 200031,China

Dongguan Innovation Institute,Guangdong Medical University,Dongguan,Guangdong 523808,China

University of Chinese Academy of Sciences,Beijing 100049,China

School of Life Science and Technology,ShanghaiTech University,Shanghai 200031,China