School of Software Engineering,Beijing Jiaotong University,Beijing 100044,China

Department of Biological Sciences,Center for Systems Biology,The University of Texas at Dallas,Richardson,TX 75080,USA

MOE Key Laboratory of Bioinformatics,BNRIST Bioinformatics Division,Department of Automation,Tsinghua University,Beijing 100084,China

Institute of Science and Technology for Brain-Inspired Intelligence,Fudan University,Shanghai 200433,China