Key Laboratory of Breeding Biotechnology and Sustainable Aquaculture(CAS),Hubei Hongshan Laboratory,Key Laboratory of Aquaculture Disease Control,Ministry of Agriculture and Rural Affairs,The Innovation Academy of Seed Design,Institute of Hydrobiology,Chinese Academy of Sciences,Wuhan,Hubei 430072,China

College of Advanced Agricultural Sciences,University of Chinese Academy of Sciences,Beijing 100049,China

College of Fisheries and Life Science,Dalian Ocean University,Dalian,Liaoning 116023,China