State Key Laboratory of Tree Genetics and Breeding,Co-Innovation Center for Sustainable Forestry in Southern China,Key Laboratory of Tree Genetics and Biotechnology of Educational Department of China,Key Laboratory of Tree Genetics and Silvicultural Sciences of Jiangsu Province,Nanjing Forestry University,Nanjing 210037,China

College of Information Science and Technology,Nanjing Forestry University,Nanjing 210037,China

Research Institute of Subtropical Forestry,Chinese Academy of Forestry,Hangzhou 311400,China

Co-Innovation Center for Sustainable Forestry in Southern China,College of Life Sciences,Nanjing Forestry University,Nanjing 210037,China

Department of Paleontology,University of Vienna,Vienna,Austria