State Key Laboratory of Biocontrol and Guangdong Provincial Key Laboratory of Plant Resources,School of Life Sciences/School of Ecology,Sun Yat-sen University,Guangzhou 510275,China

Geological Institute,Russian Academy of Sciences,Moscow 119017,Russia

Tsitsin Main Botanical Garden,Russian Academy of Sciences,Moscow 127276,Russia

Department of Biogeography,Geographical Faculty,M.V.Lomonosov Moscow State University,Moscow 119991,Russia

Borissiak Paleontological Institute,Russian Academy of Sciences,Moscow 117997,Russia

CAS Key Laboratory of Tropical Forest Ecology,Xishuangbanna Tropical Botanical Garden,Chinese Academy of Sciences,Mengla 666303,China

Natural History Museum of Guangxi,Nanning 530012,China

School of Earth Science and Resources,Chang'an University,Xi'an 710054,China