State Key Laboratory of Herbage Improvement and Grassland Agro-ecosystems,College of Ecology,Lanzhou University,Lanzhou 730000,China

College of Life Sciences,Shanxi Normal University,Taiyuan 030031,China

College of Life Sciences,Northwest Normal University,Lanzhou 730070,China

National Key Laboratory for Tropical Crop Breeding,Sanya Institute of Breeding and Multiplication,Hainan University,Sanya 572025,China

School of Tropical Agriculture and Forestry,Hainan University,Haikou 570228,China

State Key Laboratory of Plant Diversity and Specialty Crops,Institute of Botany,the Chinese Academy of Sciences,Beijing 100093,China

State Key Laboratory of Systematic and Evolutionary Botany,Institute of Botany,Chinese Academy of Sciences,Beijing 100093,China

University of Chinese Academy of Sciences,Beijing 100049,China