State Key Laboratory of Protein and Plant Gene Research,School of Life Sciences,Peking University,Beijing 100871,China

Peking-Tsinghua Center for Life Sciences,Academy for Advanced Interdisciplinary Studies,Peking University,Beijing 100871,China

Biology Department,City College of New York,City University of New York,New York 10031 USA

PhD Program in Biology,City University of New York,New York 10016 USA

Entomology Section,National Museum of Natural History,Manila 1000,Philippines