State Key Laboratory of Maize Bio-breeding,Frontiers Science Center for Molecular Design Breeding,Joint International Research Laboratory of Crop Molecular Breeding,National Maize Improvement Center,College of Agronomy and Biotechnology,China Agricultural University,Beijing 100193,China

Sanya Institute of China Agricultural University,Sanya 572025,China

Hainan Yazhou Bay Seed Laboratory,Sanya 572025,China

State Key Laboratory of Tree Genetics and Breeding/Key Laboratory of Tree Breeding and Cultivation of the State Forestry and Grass-land Administration/Research Institute of Forestry,Chinese Academy of Forestry,Beijing 100091,China

Shanghai Key Laboratory of Plant Molecular Sciences,College of Life Sciences,Shanghai Normal University,Shanghai 200234,China

College of Horticulture,South China Agricultural University,Guangzhou,510640,China

National Center for Gene Research,CAS Center for Excellence in Molecular Plant Sciences,Chinese Academy of Sciences,Shanghai 200233,China