Center for Developmental Biology,Institute of Genetics and Developmental Biology,Chinese Academy of Sciences,Beijing 100101,China

Chongqing Key Laboratory of Plant Resource Conservation and Germplasm Innovation,Integrative Science Center of Germplasm Creation Western China(Chongqing)Science City,School of Life Sciences,Southwest University,Chongqing,400715 China

Key Laboratory of Seed Innovation,Institute of Genetics and Developmental Biology,Chinese Academy of Sciences,Beijing 100101,China

Center for Molecular Agrobiology,Institute of Genetics and Developmental Biology,Chinese Academy of Sciences,Beijing 100101,China