State Key Laboratory of Genetic Evolution & Animal Models,Key Laboratory of Bioactive Peptides of Yunnan Province,KIZ-CUHK Joint Laboratory of Bioresources and Molecular Research in Common Diseases,Kunming Institute of Zoology,Chinese Academy of Sciences,Kunming 650223,China

School of Pharmaceutical Science & Yunnan Key Laboratory of Pharmacology for Natural Products,Kunming Medical University,Kunming 650500,China

University of Chinese Academy of Sciences,Beijing 100049,China

KIZ-SU Joint Laboratory of Animal Model and Drug Development,College of Pharmaceutical Sciences,Soochow University,Suzhou 215021,China