State Key Laboratory for Mineral Deposits Research,School of Earth Sciences and Engineering and Frontiers Science Center for Critical Earth Material Cycling,Nanjing University,Nanjing 210023,China

State Key Laboratory of Tibetan Plateau Earth System,Resources and Environment,Institute of Tibetan Plateau Research,Chinese Academy of Sciences,Beijing 100101,China

State Key Laboratory of Palaeobiology and Stratigraphy,Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Palaeoenvironment,Chinese Academy of Sciences,Nanjing 210008,China

College of Earth and Planetary Sciences,University of Chinese Academy of Sciences,Beijing 100049,China

Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences,Institute of Vertebrate Paleontology and Paleoanthropology,Chinese Academy of Sciences,Beijing 100044,China