State Key Laboratory of Molecular Biology,CAS Center for Excellence in Molecular Cell Science,Institute of Biochemistry and Cell Biology,Chinese Academy of Sciences,Shanghai 200031,China

Key Laboratory of Systems Health Science of Zhejiang Province,School of Life Science,Hangzhou Institute for Advanced Study,University of Chinese Academy of Sciences,Hang-zhou,Zhejiang 310024,China

Department of Neurosurgery,Children's Hospital of Fudan University,Shanghai 201102,China

School of Life Science and Technology,ShanghaiTech University,Shanghai 201210,China