Department of Neurosurgery,Huashan Hospital,Institute for Translational Brain Research,State Key Laboratory of Medical Neurobiology,MOE Frontiers Center for Brain Science,MOE Innovative Center for New Drug Development of Immune Inflammatory Diseases,Fudan University,Shanghai 200040,China

Department of Laboratory Animal Science,MOE Frontiers Center for Brain Science,Fudan University,Shanghai 200032,China

Neuroscience Research Institute and Department of Neurobiology,School of Basic Medical Sciences,Key Laboratory for Neuroscience,Ministry of Education/National Health Commission and State Key Laboratory of Natural and Biomimetic Drugs,Peking University,Beijing 100083,China

PKU-IDG/McGovern Institute for Brain Research,Peking University,Beijing 100871,China

Faculty of Medicine and Pharmaceutical Sciences,Zhejiang University,Hangzhou 310014,China

School of Medicine,Tsinghua University,Beijing 100084,China

IDG/McGovern Institute for Brain Research,Center for Life Sciences,School of Life Sciences,Peking University,Beijing 100871,China

Chinese Institute for Brain Research,Beijing 102206,China

Department of Life Sciences,National Natural Science Foundation of China,Beijing 100085,China