Department of Molecular Evolution and Plant Systematics& Herbarium(LZ),Institute of Biology,Leipzig University,Johannisallee 21-23,Leipzig D-04103,Germany

German Centre for Integrative Biodiversity Research(iDiv)Halle-Jena-Leipzig,Puschstrasse 4,Leipzig D-04103,Germany

AMAP,IRD,CIRAD,CNRS,INRAE,University of Montpellier,Montpellier,France