Institute of Special Animal and Plant Sciences,Chinese Academy of Agricultural Sciences,Changchun 130000,China

The National Key Facility for Crop Gene Resources and Genetic Improvement,Institute of Crop Sciences,Chinese Academy of Agricultural Sciences,Beijing 100081,China

College of Life Science,Nanjing Agricultural University,Nanjing 210095,China

Agronomy College/National Key Laboratory of Wheat and Maize Crop Science/CIMMYT-China(Henan)Joint Center of Wheat and Maize,Henan Agricultural University,Zhengzhou 450002,China

Department of Plant Sciences,University of California,Davis,CA 95616,USA

Crop Research Institute,Shandong Academy of Agricultural Sciences,Jinan 250100,China