State Key Laboratory of Wheat and Maize Crop Science,and Center for Crop Genome Engineering,College of Agronomy,Henan Agricultural University,Zhengzhou 450046,China

Collaborative Innovation Center of Henan Grain Crops,College of Agronomy,Henan Agricultural University,Zhengzhou 450046,China

State Key Laboratory of Crop Stress Adaptation and Improvement,College of Agronomy,Henan University,Kaifeng 475004,China

The Shennong Laboratory,Zhengzhou 450002,China