College of Life Science,Hebei Normal University,Shijiazhuang 050024,China

Dry Land Farming Institute of Hebei Academy of Agricultural and Forestry Sciences,Key Laboratory of Crop Drought Tolerance Research of Hebei Province,Hengshui 053000,China

Institute of Crop Sciences,Chinese Academy of Agricultural Sciences/State Key Laboratory of Crop Gene Resources and Breeding,Beijing 100081,China

Biotechnology Research Institute,Chinese Academy of Agricultural Sciences,Beijing 100081,China

Institute of Crop Sciences,Chinese Academy of Agricultural Sciences,State Key Laboratory of Crop Gene Resources and Breeding,Beijing 100081,China