State Key Laboratory for Biology of Plant Diseases and Insect Pests,Institute of Plant Protection,Chinese Academy of Agricultural Sciences,Beijing 100193,China

Department of Plant Pathology,China Agricultural University,Beijing 100193,China

Scientific Observing and Experimental Station of Crop Pests in Guilin,Ministry of Agriculture and Rural Affairs,Guilin 541399,China

School of Plant Protection,Hainan University,Haikou 570228,China

State Key Laboratory of Rice Biology Institute of Biotechnology,Zhejiang University,Hangzhou 310058,China

Key Laboratory of Gene Editing Technologies,Ministry of Agricultural and Rural Affairs,Sanya 572024,China