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林火扰动后次生白桦林下重引入落叶松种子萌发的影响因素

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在黑龙江省大兴安岭地区漠河市西南部林火干扰后林地,选择 5 种不同经营措施形成的林分类型——①白桦纯林,②存在落叶松幼苗更新的白桦林,③主林层为白桦、副林层为落叶松的白桦林,④落叶松-白桦混交林,⑤落叶松纯林;每种林分类型设置 4 个 60 m×60 m的试验区,每个试验区 4 角设立 4 个 20 m×20 m模拟自然落种的样地;在种子重引入试验区样地分别模拟 4 种种子与地表接触方式——直接播种(模拟种子与凋落物层接触)、去除凋落物层播种(模拟种子与土表接触)、穴状破土播种(模拟种子与穴状破土土壤接触)、条状破土播种(模拟种子与条状破土土壤接触),于 2022 年 10 月初地表积雪覆盖前播种。测定样地灌木和草本的植物种名、植株(丛)数量,测量样方凋落物厚度,调查每个播种区域落叶松种子出苗情况,计算 5 种林型灌木层和草本层物种多样性指数——马加利夫丰富度指数(Margalef richness index)、辛普森优势度指数(Simpson index)、香农-威纳多样性指数(Shannon-Wiener index)、皮卢均匀度指数(Pielou's evenness index);采用单因素方差分析法(One-Way ANOVE)、最小显著性差异法(LSD)、皮尔逊(Pearson)相关性分析法,分析播种方式(种子与地表接触方式)、林分类型、植物多样性对种子萌发率的影响。结果表明:穴状破土和条状破土显著提高种子萌发率,不同林分类型对种子萌发率的影响不显著。穴状破土播种的种子萌发率,与灌木种数量、灌木马加利夫丰富度指数、香农-威纳多样性指数、皮卢均匀度指数均呈显著正相关,与草本种数量、白桦占比呈显著负相关;条状破土播种的种子萌发率,与草本盖度、凋落物厚度、皮卢均匀度指数呈显著正相关,与草本香农-威纳多样性指数、辛普森优势度指数呈显著负相关;去除凋落物层后播种的种子萌发率,与乔木种数量呈显著正相关。
Influencing Factors the Reintroduction of Larix gmelinii Seed Germination Under Secondary Betula platyphylla Forest After Fire Disturbance
In the southwestern part of Mohe City in the Daxing'anling region of Heilongjiang Province,following fire disturb-ance,we selected five types of forest stands formed by different management practices:(1)pure Betula platyphylla forest,(2)B.platyphylla forest with the regeneration of Larix gmelinii seedlings,(3)B.platyphylla forest with a main canopy of B.platyphylla and a secondary layer of L.gmelinii,(4)mixed forest of L.gmelinii and B.platyphylla,and(5)pure L.gmelinii forest.For each forest stand type,four experimental plots of 60 m×60 m were established,with four sampling sites of 20 m×20 m set at the corners of each experimental area to simulate natural seed dispersal.In the seed reintroduction ex-perimental plots,we simulated four different seed-soil contact methods:direct seeding(simulating seed contact with the litter layer),sowing after removing the litter layer(simulating seed contact with the soil surface),hole-shaped soil break-ing seeding(simulating seed contact with hole-shaped soil breaking),and strip-shaped soil breaking seeding(simulating seed contact with strip-shaped soil breaking seeding),with sowing conducted in early October 2022 before snow cover oc-curred.We recorded the plant species and the number of individuals(clumps)for shrubs and herbaceous plants in the sampling plots,measured the thickness of the litter layer,and surveyed the germination status of L.gmelinii seeds in each sowing area.We calculated the species diversity indices for the shrub and herb layers of the five forest types,including the Margalef richness index,Simpson index,Shannon-Wiener diversity index,and Pielou's evenness index.One-Way ANO-VA,Least Significant Difference(LSD),and Pearson correlation analyses were employed to assess the effects of sowing method(seed-soil contact method),forest type,and plant diversity on seed germination rates.The results showed that the hole-shaped and strip-shaped soil breaking significantly increased seed germination rates,whereas the influence of different forest stand types on seed germination rates was not significant.The seed germination rate for hole-shaped and strip-shaped soil breaking showed a significant positive correlation with the number of shrub species,the Margalef richness index of shrubs,the Shannon-Wiener diversity index,and the Pielou's evenness index,while displaying a significant negative cor-relation with the number of herb species and the proportion of B.platyphylla.For strip soil breaking sowing,the seed ger-mination rate was significantly positively correlated with the coverage of herbaceous plants,litter thickness,and Pielou's evenness index,while being negatively correlated with the herbaceous Shannon-Wiener diversity index and Simpson index.The seed germination rate for sowing after removing the litter layer was significantly positively correlated with the number of tree species.

Burned areaForest regenerationBetula platyphyllaLarix gmeliniiSeed germinationGreat Xing'an Mountain

唐慧敏、宋晓倩、金鹭、施宇、关鑫、陈嘉琦、张衷华、唐中华

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东北林业大学,哈尔滨,150040

火烧迹地 森林更新 白桦 落叶松 种子萌发 大兴安岭

2025

东北林业大学学报
东北林业大学

东北林业大学学报

北大核心
影响因子:0.74
ISSN:1000-5382
年,卷(期):2025.53(2)