Department of Biochemistry and Molecular Biology,School of Basic Medicine,Tongji Medical College,Huazhong University of Science and Technology,Wuhan 430030,China

Cell Architecture Research Center,Huazhong University of Science and Technology,Wuhan 430030,China

State Key Laboratory for Diagnosis and Treatment of Severe Zoonotic Infectious Diseases,Huazhong University of Science and Technology,Wuhan 430030,China

Shenzhen Bay Laboratory,Shenzhen 518132,China