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川西高山森林生态系统林下生物量及其随林窗的变化特征

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作为森林生态系统的重要组成部分,林下植被及其残体的分布受到林冠层的影响,但迄今有关林窗对林下植被和残体生物量的影响尚无研究报道。于2013年8月2日至20日,以海拔3600 m的川西岷江冷杉原始林林下植被为研究对象,根据区域内的坡向和林分组成等因素设置3个100 m×100 m的典型样地,调查其生物量及其随林窗的变化特征。在每个样地内选择3个大林窗,在林窗、林缘和林下分别设置3个20 m×20 m的样方,调查粗木质残体长度或高度、大小头直径、枯立木记录胸径、腐烂等级等;在林窗、林缘和林下分别设置3个5 m×5 m的样方,采用“收获法”收集样方内直径在2.5~10 cm之间的细木质残体和灌木生物量;在林窗、林缘和林下分别设置3个1 m×1 m的样方来调查凋落物储量和草本生物量;在1 m×1 m的样方内随机选择1个20 cm×20 cm的小样方来调查地被植物生物量。结果表明,(1)川西高山森林生态系统总生物量为72.75 t·hm-2,其中林下生物量为67.92 t·hm-2,占生态系统生物量的95.17%。活体植被以灌木为主,其生物量为9.81 t·hm-2;残体部分以粗木质残体为主,其储量为53.00 t·hm-2;(2)林窗对灌木、草本、地被植物的影响各不相同,且不同物种的灌木生物量表现出不同的分布规律;草本生物量表现出明显的“边缘效应”,在林缘显著高于林下;林窗和林缘的地被植物生物量相对较低;(3)粗木质残体储量从林下到林窗呈现减小的趋势,但总体储量仍然较大,林窗和林缘的细木质残体储量高于林下。这些结果为认识高山森林生态系统林下生物量及其格局,以及林窗在森林生态系统的重要作用提供了基础理论依据。
Understory biomass and its characteristics as affected by forest gap in the alpine forest ecosystem in west Sichuan
Understory vegetation and its debris, as an important component in the forest ecosystem, can be greatly affected by forest canopy. As yet, little information has been available on the effects of forest gaps on understory plant and its debris biomass. In order to understand the changes of understory plant and its debris biomass as affected by forest gaps, therefore, three 100 m×100 m plots were sampled in a Minjiang fir (Abies faxoniana) primary forest ecosystem around 3 600 m altitude in western Sichuan in August, 2013. Coarse woody debris with length/height, basic diameter, diameter at breast height and decay class were investigated in each of three 20 m×20 m sampling quadrats in forest gap center, gap edge and closed canopy, respectively. Fine woody debris and shrub biomass were investigated in each of three 5 m×5 m sampling quadrats in forest gap, gap edge and closed canopy, respectively. Moreover, non-woody litter and herb biomass were harvested in each of three 1 m×1 m sampling quadrats in forest gap, forest edge and closed canopy, respectively. Each of 20 cm×20 cm sampling quadrat was randomly selected to harvest groundcover vegetation biomass in the above 1 m×1 m quadrat. The results indicated that:(1) the total biomass was 72.75 t·hm-2 in the sampled alpine forest ecosystem, and understory biomass was 67.92 t·hm-2 which accounted for 95.17%of the total biomass in whole ecosystem. Shrub biomass was 9.81 t·hm-2 and occupied the majority proportion of living vegetation, but coarse woody debris biomass was 53.00 t·hm-2 and occupied the majority of dead debris. (2) Shrub, herb and groundcover vegetation biomass showed different responses to forest gap. Shrub biomass was significantly higher in forest edge than that in closed canopy, although the biomass of groundcover vegetation was relatively lower in forest gap and forest edge compared with that in closed canopy. (3) Coarse woody debris biomass showed a decrease tendency from closed canopy to gap center, but the storage of fine woody debris in gap center and forest edge was larger than that in closed canopy. These results provide efficient foundation in understanding the understory biomass and its pattern in these alpine forest ecosystems as well as understanding the important role of gaps.

alpine forestunderstory vegetationbiomassforest gap

肖洒、吴福忠、杨万勤、常晨辉、李俊、王滨、曹艺

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四川农业大学生态林业研究所,林业生态工程重点实验室,高山森林生态系统定位研究站,四川 成都 611130

高山森林 林下植被 生物量 林窗

国家自然科学基金国家自然科学基金国家“十二五”科技支撑计划四川省杰出青年学术与技术带头人培育项目四川省杰出青年学术与技术带头人培育项目中国博士后科学基金特别资助

31170423312704982011BAC09B052012JQ00082012JQ00592012T50782

2014

生态环境学报
广东省生态环境与土壤研究所 广东省土壤学会

生态环境学报

CSTPCDCSCD北大核心
影响因子:1.608
ISSN:1674-5906
年,卷(期):2014.(9)
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