Hubei Hongshan Laboratory,Wuhan 430070,Hubei,China

MOA Key Laboratory of Crop Ecophysiology and Farming System in the Middle Reaches of the Yangtze River,The Center of Crop Nanobiotechnology,College of Plant Science&Technology,Huazhong Agricultural University,Wuhan 430070,Hubei,China

School of Science,Hawkesbury Institute for the Environment,Western Sydney University,Penrith,NSW 2751,Australia